During the German occupation of France, Shosanna Dreyfus saw the execution of his family to hands of the colonel nazi Hans Landa. Shosanna escape to Paris, where she obtained a new identity as owner and director of a cinema.
In another place of Europe, the lieutenant Aldo Raine organizes a group of Jewish soldiers to attack to concrete aims. Known by the enemy as "The Basterds", Raine's troops join the German actress Bridget Von Hammersmark, a secret agent who works for the allies, in order to do a mission that will make fall the leaders of the Third Reich. The destiny wants that they meet under the marquee of a cinema where Shosanna waits to take revenge.
In the cinema opened a nazi film and Hitler goes to see this film. While the leaders of the Third Reich are into the cinema Shosanna burns it. The troops of Aldo Rayne go in the cinema and kill Hitler. Only survive Aldo Raine who returns to his homeland with the leader of SS as a prisoner.
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